Rabbit coat genetics

Prior to domestication, rabbits were all of the agouti or wild-type color pattern, characterized by alternating bands of black and red color on the hair shaft.

Major coat color genes
Most of the genes that affect coat colors on rabbits are located on four pairs of chromosomes. These genes have been categorized into a system referred to as the coat color series and are labeled A to E (agouti, brown, color, dilution, and extension).

Coat pattern genes
Other genes other than the major A to E coat color series exist that also exert pronounced effects on the coat color pattern.

En series: Broken
Charlies have two En/En genes, normal broken rabbits have En/en genes, while normal solid colored rabbits have en/en genes.